We thank the Cryo-EM Center at University of Science and Technology of China for the support of cryo-EM data collection. This work was funded by the National Natural Science Foundation of China (21825703 and 21927814 to C.T., 31971152 and 22277114 to P.S., 22307113 to S.L.), the National Key R&D Program of China (2021YFA1200104 to S.L., 2022YFC3400500 to C.T.), the Strategic Priority Research Program of Chinese Academy of Sciences (XDB37000000 to C.T.), and the Natural Science Foundation of Anhui Province (2108085J16 to P.S.).