1 Center for Theoretical Biological Physics, Northeastern University, Boston, MA, United States of America

2 Institut für Theoretische Physik, Leopold-Franzens-Universität Innsbruck, Technikerstrasse 21a, A-6020 Innsbruck, Austria

3 Institute of Mathematics and Statistics, University of Tartu, Tartu, Estonia

4 Center for Complexity and Biosystems, Department of Environmental Science and Policy, University of Milan, via Celoria 10, 20133 Milano, Italy

5 Center for Complexity and Biosystems, Department of Physics, University of Milan, via Celoria 16, 20133 Milano, Italy

6 CNR—Consiglio Nazionale delle Ricerche, Istituto di Chimica della Materia Condensata e di Tecnologie per l'Energia, Lecco, Italy

7 CNR—Consiglio Nazionale delle Ricerche, Istituto di Biofisica, via Via De Marini 6, 16149 Genova, Italy

8 Department of Physics and Bioengineering, Northeastern University, Boston, MA, United States of America