These authors contributed equally: Cory T. Miller, Weizhe Hong.

Cortical Systems & Behavior Lab, University of California, San Diego, La Jolla, CA, USA

Cory T. Miller

Department of Biology, Stanford University, Stanford, CA, USA

Xiaoke Chen

Department of Molecular, Cellular & Developmental Biology, University of Colorado Boulder, Boulder, CO, USA

Zoe R. Donaldson

Department of Psychology & Neuroscience, University of Colorado Boulder, Boulder, CO, USA

Zoe R. Donaldson

Department of Psychology, Columbia University, New York, NY, USA

Bianca Jones Marlin

Department of Neuroscience, Columbia University, New York, NY, USA

Bianca Jones Marlin

Department of Molecular and Cell Biology, Howard Hughes Medical Institute, University of California, Berkeley, Berkeley, CA, USA

Doris Y. Tsao

Massachusetts General Hospital, Harvard Medical School, Boston, MA, USA

Ziv M. Williams

Department of Neurobiology, University of Utah, Salt Lake City, UT, USA

Moriel Zelikowsky

Allen Institute for Brain Science, Seattle, WA, USA

Hongkui Zeng

Department of Neurobiology, University of California, Los Angeles, Los Angeles, CA, USA

Weizhe Hong

Department of Biological Chemistry, University of California, Los Angeles, Los Angeles, CA, USA

Weizhe Hong