These authors contributed equally: Zijie Feng, Xin He

Department of Cancer Biology, Abramson Family Cancer Research Institute, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA, USA

Zijie Feng, Xin He, Xuyao Zhang, Yuan Wu, Bowen Xing, Alison Knowles, Qiaonan Shan, Samuel Miller, Taylor Hojnacki, Jian Ma, Bryson W. Katona & Xianxin Hua

Division of Gastroenterology and Hepatology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA, USA

Bryson W. Katona & David C. Metz

Department of Radiology, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA, USA

Terence P. F. Gade

Center for Cellular Immunotherapies, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA, USA

Don L. Siegel & Carl H. June

Department of Pathology and Laboratory Medicine, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA, USA

Don L. Siegel & Carl H. June

I. Department of Medicine, University Medical Center Hamburg-Eppendorf, Hamburg, Germany

Jörg Schrader