Division of Quantitative Biology, Okazaki Institute for Integrative Bioscience, National Institute for Basic Biology, National Institutes of Natural Sciences, Aichi, 444-8787 Japan

Quantitative Biology Research Group, Exploratory Research Center on Life and Living Systems (ExCELLS), National Institutes of Natural Sciences, Aichi, 444-8787 Japan

Department of Basic Biology, School of Life Science, SOKENDAI (The Graduate University for Advanced Studies), Aichi, 444-8787 Japan